# Lab Alumni Sort ![Headshot of L Lai](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/philip_lai_updated_9.14.22_resized.jpg?itok=Iin-zoV1) **L. Fahn-Lai** Graduate Student 2015-2021, **PhD 2021** L. studies the tapestry of muscle and bone that makes up the shoulder, and how its structure and function evolved along the branch of the vertebrate tree of life that led to modern-day mammals. Their work combines CT imaging, high-speed X-ray videography, machine learning, and old-fashioned probe and scalpel dissection to produce new insights into the inner workings of ancient anatomical structures. Following their PhD, L. is pursuing a career in Data Design Science. [L Fahn-Lai Website](https://fahnl.ai/) ![Headshot of Kari Taylor-Burt](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/kari_taylor-burt.jpg?itok=mD8rOclK) **Kari Taylor-Burt** Graduate Student 2013-2020, **PhD 2020** Kari's interests focus on how environmental demands influence the biomechanics of animal movement, musculoskeletal structure, and physiology. Kari is currently an Assistant Professor of Biology at Mt. Saint Mary University in Pennsylvania working on invertebrate obliquely striated muscle. [Kari Taylor-Burt Website](https://karitaylorburt.com/)​​​​​​​ [Mount Saint Mary's Univ Website](https://directory.msmary.edu/people/kari-taylor-burt.html) ![Headshot of Brianna McHorse](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/brianna_mchorseprofile.jpg?itok=MM2k3M_K) **Brianna McHorse** Graduate Student 2013-2019, **PhD 2019** Research Interests: terrestrial locomotion, functional morphology, evolutionary biomechanics, paleobiology, musculoskeletal modeling, equid evolution. Following her PhD, Brianna is pursuing a career in Data Science. ![Allison Arnold sitting in front of computer](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/allison_arnold.jpg?itok=qqUZFy_K) **Allison Arnold-Rife, PhD** Research Associate, 2005-2019 Allison's research Interests include musculoskeletal modeling and simulation of human and animal movement; experimental validation of muscle models; applications of models in comparative biomechanics, pediatric orthopedics, rehabilitation, and sports. Allison currently works as an independent scientist, collaborating in the Boston, MA area. [Allison Arnold-Rife Profile](https://www.linkedin.com/in/asarnoldrife/) ![Headshot of Chris Tijs](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/christijs_0.jpg?itok=QK9oxhFl) **Chris Tijs, PhD** NIH Post-doc 2015-2017 Research Interests: biomechanics, muscle and tendon mechanics, connective tissue, myofascial force transmission. Following his Post-doc at Harvard's CFS, Chris returned to the Netherlands to pursue a career as a Data Analyst, working for Netherlands Translink. [Chris Tijs Profile](https://nl.linkedin.com/in/christijs) ![Partha Bhagavatula holding pigeon](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/parthacfswpigeon2.jpg?itok=C8KF0zYa) **Partha Bhagavatula, PhD** ONR Post-doc 2014-2016 Research Interests: avian vision, flight control in birds, animal navigation and GNSS (Global Navigation Satellite system) tools for animal tracking. partha.bhagavatula\[at\]gmail.com Following his Post-doc at Harvard's CFS, Partha has pursued a series of post-docs in Singapore, Minnesota and Houston, TX. ![Glenna](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/glenna_clifton.jpg?itok=8ZkD8Ju2) **Glenna Clifton** Graduate Student 2000-2017, **PhD 2017** Research Interests: underwater force production, predator-prey interactions of diving birds, running on water by Western and Clark's grebes, swing limb muscle control. Following her PhD, Glenna was a post-doc in Bioengineering at UCSD, and is currently an Assistant Professor of Biology at the University of Portland. glenna.clifton\[at\]gmail.com [Glenna Clifton Website](http://www.glennaclifton.com/) ![Headshot of Jen Carr](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/jen_carr.jpg?itok=SSARFBMd) **Jennifer Carr, PhD** Preceptor and Instructor, Department of Organismic & Evolutionary Biology, 2009-2016 Research interests: terrestrial locomotion, muscle morphology, evolutionary morphology, muscle function during terrestrial locomotion Jen is currently a Lecturer and Undergraduate Lab Coordinator at Salem St. University ![Headshot of Talia Moore](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/talia_moore.jpg?itok=PteHYjXL) **Talia Y. Moore** Graduate Student 2000-2016, **PhD 2016** Research interests: jerboas, kangaroo rats, hopping mice, bipedal desert rodents, terrestrial locomotion and associated morphological adaptations, granular substrates, phylogenetic methods, inertial steering. Following her PhD, Talia was a post-doc in Bioengineering at the University of MIchigan, and is currently an Assistant Professor of Robotics in the Mechanical Engineering department at the University of MIchigan. taliaym\[at\]gmail.com [Talia Moore Faculty Profile](https://robotics.umich.edu/profile/talia-moore/) ![Headshot of Carolyn Eng](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/carolyneng2.jpg?itok=7qf-SKUO) **Carolyn Eng** Graduate Student 2008-2014, **PhD 2014** NIH Post-doc 2014-15 Research Interests: muscle architecture and biomechanics of movement, musculoskeletal modeling, myofascial and myotendinous biomechanics. Following her NRSA Post-doc at Brown University and a post-doc at Yale University, Carolyn opted to pursue a career as a Research Project Manager and is now a Senior Scientific Project and Alliance Manager at the Allen Institute for Neural Dynamics, Seattle, WA. carolyneng\[at\]gmail.com [Carolyn Eng Profile](https://www.linkedin.com/in/carolyn-eng) ![Ivo Ros with a cockatiel perched on finger](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/ivoros_cfs.jpg?itok=NwzZGHH4) **Ivo Ros** Graduate Student, 2008-2013, **PhD 2013**; ONR Post-doctoral fellow 2014-15 Research Interests: aerodynamics and visual control of maneuvering and turning flight in birds and flies. Following his PhD and an ONR Post-doctoral fellowship at Harvard, Ivo continues his academic research as a Research Associate at the California Institue of Techology, studying the neurobiology of flight control in fruit flies (*D. melanogater*), analyzing their neural circuit dynamics based on functional imaging via optogenetics and 2-photon microscopy. ivo.ros\[at\]gmail.com [Ivo Ros Profile](https://www.linkedin.com/in/ivo-ros-a17b53a2) [Dickinson Lab - Cal Tech](https://dickinsonlab.caltech.edu/people/ivo-ros/) ![Headshot of C. David Williams](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/davewilliams_cfs.jpg?itok=M-jKUFbp) **C. David Williams, PhD** NSF Post-doctoral Fellow 2012-14 Research Interests: molecular analysis of sarcomere force-length behavior, flight stability and maneuvering in birds, and currently BIG data science. Dave returned to the Univ. of Washington as a mathematical biophysicist at the Allen Institute, and is currently a Data & Modelling Scientist at Microsoft. cdave\[at\]uw.edu [C. David Williams Website](http://charlesdavidwilliams.com/) ![Headshot of Natalie Holt](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/natalieholtcfs.jpg?itok=l9yqtG8L) **Natalie Holt, PhD** NIH Post-doctoral Researcher, 2011 - 2013 Research Interests: comparative muscle physiology, energetics and biomechanics of animal locomotion After a Post-doc at UC, Irvine, Natalie was an Assistant Professor of Biology at Northern Arizona Univeristy, and is now an Assistant Professor of Evolution, Ecology & Organismal Biology at UC, Riverside. natalie.c.holt\[at\]gmail.com [Holt Lab Website](https://theholtlab.weebly.com/) ![Maria de Boef Miara standing among large rocks](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/de_boef_miara_0.jpg?itok=8AfJvgtb) **Maria de Boef Miara, PhD** 2007 - 2012 NIH Post-doctoral fellow Research Interests: terrestrial locomotion, musculoskeletal morphology, bone microstructure, evolution and development. Maria is currently and Associate Professor of Biology, teaching at Brandeis University [Maria de Boef Miara Profile](https://www.brandeis.edu/neuroscience/faculty/miara-maria.html) ![Headshot of Huai-Ti Lin](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/lin_0.jpg?itok=10DO6LPv) **Huai-Ti Lin, PhD** ONR Postdoctoral Researcher, 2011 - 2011 Research Interests: avian and insect flight and navigation through cluttered environments; soft-bodied robotics; predator-prey pursuit in dragonflies; flight stabiligy and biorobotics. After completing a Research Associate fellowship at Janelia Farms, HHMI, Huai-Ti moved to a faculty position at Imperial College, London, where he is now Associate Professor of Bioengineering. h.lin\[at\]imperial.ac.uk [Huai-Ti Lin Profile](https://www.imperial.ac.uk/people/h.lin) [ Huai-Ti Lin Lab Website](https://htlinlab.com/) ![Headshot of Carlos Moreno](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/moreno.jpg?itok=ohpCrHyJ) **Carlos Moreno** Graduate Student, 2004 - 2010, **PhD 2010** Academic Research Interests: skeletal biomechanics and biomechanics of goat turning during steady locomotion. Carlos is currently a Senior Software Engineer, Stem Disintermedia Inc. [Carlos Moreno Profile](https://www.linkedin.com/in/carlos-moreno-52b3b347) ![Headshot of Angie Berg Robertson](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/robertson.jpg?itok=xagpEXSp) **Angie Berg Robertson** Graduate Student, 2004 - 2010, **PhD** Academic Research Interests: avian flight biomechanics and aerodynamics. Currently: pursuing family life in Phoenix, AZ ![Chris Richards standy by lab equipment](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/richards.jpg?itok=UwxcujQq) **Chris Richards** Graduate Student, 2003 - 2009, **PhD** Research Interests: biorobotics, modeling and experimental muscle biomechanics and locomotor performance in swimming frogs Following his PhD, Chris was a Junior Fellow at the Rowland Institute, Harvard University. Chris then took a research post-doctoral fellowship in paleorobotics at the Royal Veterinary College, UK. Chris is now an Associate Professor of Bioengineering at Imperial College, London. [Chris Richards Profile](https://profiles.imperial.ac.uk/c.richards1) ![Headshot of Edwin Yoo](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/yoo.jpg?itok=hz173qzo) **Edwin Yoo** Graduate Student: 2006-2009, **M.A.** (Harvard, OEB) Research: climbing mechanics in goats, horn impacts during head butting Currently: High School Science Teacher ![Headshot of David Lentink](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/lentink.jpg?itok=IvvOIQsj) **David Lentink, PhD** NSF Postdoctoral Researcher, 2009-2010 Research Interests: aerodynamics of bird flight; wing morphing linked to maneuvering in swifts; stability during avian flight maneuvering. Currently: Associate Professor of Mechanical Engineering, University Groningen, Netherlands [David Lentink Profile](https://www.rug.nl/staff/d.lentink/?lang=en) ![Headshot of Timothy Higham](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/tim_higham.jpg?itok=5iJlL8Tu) **Timothy E. Higham, PhD** NIH Postdoctoral Researcher, 2006-2008 Research Interests: vertebrate locomotion and feeding, muscle biomechanics, functional morphology Currently: Professor of Evolution, Ecology and Organismal Biology, UC Riverside [Timothy Higham Profile](https://profiles.ucr.edu/app/home/profile/thigham) ![Andrew Carroll kneeling and holding dog](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/andrew_carroll.jpg?itok=qMBLI0iL) **Andrew Carroll, PhD** Postdoctoral Researcher Research: vertebrate locomotion and feeding, muscle physiology, functional morphology Assistant Professor, Dept. of Biology, Evansville College Deceased ![David Lee on one knee with dog](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/david_lee.jpg?itok=h3uPca3Z) **David V. Lee, PhD** DARPA Postdoctoral Researcher, 2003-2006 Research Interests: comparative biomechanics of terrestrial locomotion; XROMM and biorobotics. Currently: Associate Professor, School of Life Sciences, University of Nevada, Las Vegas [David Lee Profile](http://www.unlv.edu/people/david-lee) ![Polly McGuigan standing in front of CFS sign](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/polly_mcguigan.jpg?itok=FvDtFBOC) **M. Polly McGuigan, PhD.** Postdoctoral Researcher, 2003-2005 Research Interests: muscle-tendon mechanics, muscle modulation with varying locomotor tasks; aging effects on muscle and locomotor function Currently: Senior Lecturer and Department Head, Applied Biomechanics, Department for Health, University of Bath, UK [Polly McGuigan Profile](http://www.bath.ac.uk/health/staff/polly-mcguigan/) ![Monica Daley with black and white dog](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/monica_daley.jpg?itok=BjMZ8YE_) **Monica A. Daley** Graduate Student, 2000-2006, **PhD** Research Interests: neuromuscular control of locomotion, perturbation recovery, locomotor biomechanics and energetics, biorobotics. Currently: Professor of Biology, School of Biological Sciences, UC, Irvine [Monica Daley Profile](https://neuromechanics.bio.uci.edu/lab-members/) ![Headshot of Russell Main](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/russ_main.jpg?itok=EB31PGhq) **Russell P. Main** Graduate Student, 2000-2006, **PhD** Research Interests: skeletal biomechanics, bone remodeling, bone microstructure and geometry, in vivo patterns of bone strain. Associate Professor, Basic Medical Sciences (Veterinary Medicine) & Weldon School of Bioengineering, Purdue University Deceased (along with fellow friend and CFS mascot "Eduardo the Emu") ![Craig McGowan holding a wallaby](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/craig_mcgowan.jpg?itok=VEusvXM0) **Craig P. McGowan** Graduate Student, 2000-2006, **PhD** Research Interests: comparative biomechanics, neuromuscular control of locomotion, musculoskeletal modeling and evolution. Currently: Associate Professor, Dept. of Integrative Anatomical Sciences, Keck School of Medicine, USC [Craig McGowan Profile](https://keck.usc.edu/faculty-search/craig-mcgowan/) ![Headshot of Ty Hedrick](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/ty_hedrick.jpg?itok=RKYN6fWR) **Ty Hedrick** Graduate Student, 1999-2004, **PhD** Research Interests: aerodynamics and control of flight, flight stability and maneuvering; 3D kinematics of flight and field studies of flight. Ty has pioneered open-source software tools to study the 3D kinematics of animal movement under a variety of conditions. Currently: Professor of Biology, University of North Carolina at Chapel Hill [Hedrick Lab Website](https://biomech.web.unc.edu/) ![Jim Usherwood facing an emu](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/jusherwood.jpg?itok=TvtaaUOt) **Jim Usherwood, PhD** Postdoctoral Researcher, 2002-2004 Research Interests: comparative biomechanics of terrestrial and aerial locomotion; remote monitoring of flight performance in birds. Currently: Wellcome Trust Senior Fellow and Professor, Structure and Motion Laboratory, Royal Veterinary College, UK [Jim Usherwood Profile](https://www.rvc.ac.uk/about/our-people/james-usherwood) ![Headshot of Anna Ahn](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/anna_ahn.jpg?itok=RTlwPWH_) **Anna Ahn, PhD** NIH Postdoctoral Researcher, 2001- 2003 Research Interests: neuromuscular control and biomechanics of locomotion, muscle mechanics. Currently: Professor of Biology, Harvey Mudd College [Anna Ahn Profile](https://www.hmc.edu/biology/faculty-staff/anna-n-ahn/) ![Headshot of Gary Gillis](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/gillis.jpg?itok=PdT5JYRK) **Gary B. Gillis, PhD** NRSA NIH Postdoctoral Researcher, 1998- 2002 Research Interests: biomechanics, neuromuscular control of locomotion; evolution of vertebrate musculoskeletal systems Currently: Norman and Emma Harris Foundation Professor of Biology, Mount Holyoke College [Gary Gillis Profile](https://www.mtholyoke.edu/directory/faculty-staff/gary-gillis) ![Headshot of Barbekka Hurtt](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/barb_hurtt.jpg?itok=zG3BmdVP) **Barbekka Hurtt, PhD** NIH Postdoctoral Researcher Research: muscle function and energetics of exercise ![Profile headshot of Bret Tobalske](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/brett_tobalske.jpg?itok=qUa9rfpc) **Bret W. Tobalske, PhD** Postdoctoral Researcher, 1998- 1999 Research Interests: aerodynamics of bird flight, flight ecology and neuromuscular physiology linked to flight performance. Currently: Professor of Biology, Division of Biological Sciences, and Director of the Field Research Station, University of Montana [University of Montana Flight Lab](https://www.umt.edu/flight-lab/default.php) ![Headshot of Peter Weyand](/sites/g/files/omnuum6301/files/styles/hwp_1_1__360x360_scale/public/biewener/files/peter_weyand.jpg?itok=wmLZDjN6) **Peter G. Weyand, PhD** Research Associate 1995-1998 Research Interests: exercise physiology, human locomotion Currently: Professor & Chair, Director Locomotor Performance Laboratory, Texas Christian University [Peter Weyand Profile](https://www.tcu.edu/directory/profile/peter-weyand.php) **Past Doctoral Students (University of Chicago, Department of Organismal Biology & Anatomy)** **Carolyn Jaslow, PhD 1987** (Professor, Rhodes College) **Sharon Swartz, PhD 1988** (Professor, Brown University) **John Bertram, PhD 1988** (Professor, Univ. of Calgary) **Larry Frolich, PhD 1991** **Matthew Carrano, PhD 1998** (Curator, Smithsonian HMNH) (joint w/ J Hopson) **Rick Blob, PhD 1998** (Professor, Clemson University) (joint w/ J Hopson) **Nora Espinoza, PhD 2000** (Adjunct Assoc. Professor, Clemson University)